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D. campyloptera resources
Descriptive
Connecticut Botanical SocietyConnecticut Botanical Society
Distribution
North AmericaNorth America
Flora
Flora of North AmericaFlora of North America
Herbarium
lower bladelower blade
Monograph
Hoshizaki and WilsonHoshizaki and Wilson
Photo
bladeblade
lowest pinnaelowest pinnae

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Dryopteris campyloptera (Kunze) Clarkson

Mountain wood fern

Etymology Latin: campylo means curved + ptera, from the Greek pteron meaning wing
Description Rhizome: erect.
Frond: 60 cm high by 25 cm wide, deciduous, monomorphic, blade/stipe ratio: 2:1.
Stipe: grooved, green, light-brown scales at the base, vascular bundles: 5 in a c-shaped pattern.
Blade: 3-pinnate-pinnatifid, deltate-ovate, herbaceous, linear to ovate scales below, extending to costa, absent above.
Pinnae: anadromous, in plane of blade, lanceolate; basal pinnae deltate to broadly lanceolate, not reduced; pinnules basiscopic pinnule of lowest pinnae very long, basal pinnules equal to adjacent pinnules, basal basiscopic pinnule much longer and 2 times width of acroscopic one; costae grooved above, continuous from rachis to costae; margins margins toothed, teeth spine-tipped.; veins free, forked.
Sori: round, in 1 row between midrib and margin, indusium: reniform, at a sinus, sporangia: brownish, maturity: midsummer.
Culture Habitat: cool, moist woods at increasing elevation southward. Distribution: eastern North America. Hardy to -30C, USDA Zone 4.
Synonyms
Aspidium campylopterum Kunze
Dryopteris austriaca (Jacquin) Schinz & Thellung
Dryopteris dilatata ssp. americana (Fisch.) Hulten
Dryopteris spinulosa (Swartz) Watt var. americana (Fischer ex Kunze) Fernald
Dryopteris spinulosa var. concordiana (Davenport) Eastman
Dryopteris campyloptera
Dryopteris campyloptera.  Illustration from The Cultivated Species of the Fern Genus Dryopteris in the United States, Barbara Joe Hoshizaki and Kenneth A. Wilson, American Fern Journal, 89, 1, (1999), with permission.
Notes
Spinulose woodferns Pinnae in plane of blade, lanceolate; basal pinnae deltate to broadly lanceolate, not reduced, basal pinnules equal to adjacent pinnules, basal basiscopic pinnule much longer and 2 times width of basal acroscopic pinnule (only in this species)
Dryopteris campyloptera
Dryopteris campyloptera. Five vascular bundles at both the base (below) and the top of the stipe. The groove on the upper surface of the stipe becomes more pronounced as one moves up.  Drawing from Ferns of Northeastern United States, Farida A. Wiley, 1936.
Dryopteris campyloptera
Dryopteris campyloptera. lowest pinna, tripinnate at base, strongly anadromic, scales extend to costa.  Illustration by V. Fulford from Ferns and Fern Allies of Canada, William J. Cody and Donald M. Britton, 1989, Agriculture Canada, used with permission.
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