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Dryopteris arguta (Kaulfuss) Maxon

Coastal wood fern

Etymology Latin: arguta means sharply toothed or notched.
Description Rhizome: short-creeping.
Frond: 60 cm high by 20 cm wide, evergreen, monomorphic, blade/stipe ratio: 2:1 to 3:1.
Stipe: grooved, scattered, light brown or reddish-brown scales, vascular bundles: 3-7 in a c-shaped pattern.
Blade: pinnate-pinnatifid to 2-pinnate at base, lanceolate, widest and truncate at the base, herbaceous or papery, glandular.
Pinnae: 17 to 18 pair, subopposite; pinnules oblong, about 1 cm; costae grooved above, continuous from rachis to costae; narrow, lanceolate scales; margins serrate to lobed, decidedly spiny; veins free, running to teeth tips.
Sori: round, medial, indusium: reniform, pale greenish yellow, at a sinus, sporangia: brownish-gray.
Culture Habitat: open, wooded slopes. Distribution: coastal British Columbia to California, Baja California, disjunct in Arizona. Hardy to -15C, USDA Zone 7; however, the majority of selections are limited to Zone 8.
Distinctive Characteristics narrow, lanceolate scales on costa; very spinulose Dryopteris
Synonyms
Aspidium argutum Kaulfuss
Lastrea arguta Brackenr.
Nephrodium argutum Diels
Thelypteris arguta Moxley
Dryopteris arguta
Dryopteris arguta.  Illustration from The Cultivated Species of the Fern Genus Dryopteris in the United States, Barbara Joe Hoshizaki and Kenneth A. Wilson, American Fern Journal, 89, 1, (1999), with permission.
Dryopteris arguta
Dryopteris arguta. a) frond, widest at the base; b) fertile pinnule, spiny.  Illustration by V. Fulford from Ferns and Fern Allies of Canada, William J. Cody and Donald M. Britton, 1989, Agriculture Canada, used with permission.
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