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Lygodium palmatum (Bernhardi) Swartz

Hartford fern, American climbing fern

Etymology Palmatum means palmately shaped.
Description Rhizome: creeping, branching, stipes 1-4 cm apart, short, blackish or red hairs.
Frond: 300 cm high , evergreen, but the fertile pinnae dying back in winter, monomorphic, blade/stipe ratio: 30:1.
Stipe: twining, branching, shining black, shading to brown and green above, hairy at base, vascular bundles: 1, a simple cylinder.
Blade: 2-pinnate, twining rachis, palmate or pinnate pinnae, transparent hairs on lower surface; a second form in northern areas is glabrous.
Pinnae: alternate, sterile pinnae on 1-2 cm stalks, a pair of pinnules very broadly ovate, deeply and palmately 3-7-lobed, to 6 cm; fertile: contracted, always at the end of the frond, irregularly, palmately 3-6-forked or lobed; costae sometimes ridged or winged; segments sterile: ultimate lobes triangular-elongate to oblong; margins entire; veins free, forking.
Sori: , indusium: false, scale-like, opening along a longitudinal slit, sporangia: in 2 rows, covering the surface of the ultimate segments, maturity: winter, short viability.
Dimensionality: pinnae held horizontally.
Culture Habitat: margins of woods, thickets, and bogs in humus-rich, acid soils, not in shade. Distribution: eastern United States, rare except in Kentucky, Tennessee. Hardy to -35�C, USDA Zone 3.
Distinctive Characteristics twining habit, palmate sterile segments
Synonyms
Gisopteris palmata Bernhardi
Ramondia palmata Bosc
Lygodium palmatum
Lygodium palmatum. Frond, sterile and fertile segments. �Illustration by Edgar Paulton, from How to Know the Ferns and Fern Allies, John T. Mickel, � 1979 Wm. C. Brown Co.
Notes
Spore Maturity Spore collected in the fall fails to germinate, while winter collection is successful -- J. Mickel, personal communication.

Lygodium palmatum. Habit. �Web pages of the Connecticut Botanical Society, with permission.
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