Etymology
Latin: campylo means curved + ptera, from the Greek pteron meaning wing
Description
Rhizome: erect.
Frond: 60 cm high by 25 cm wide, deciduous, monomorphic, blade/stipe ratio: 2:1.
Stipe: grooved, green, light-brown scales at the base, vascular bundles: 5 in a c-shaped pattern.
Blade: 3-pinnate-pinnatifid, deltate-ovate, herbaceous, linear to ovate scales below, extending to costa, absent above.
Pinnae: anadromous, in plane of blade, lanceolate; basal pinnae deltate to broadly lanceolate, not reduced; pinnules basiscopic pinnule of lowest pinnae very long, basal pinnules equal to adjacent pinnules, basal basiscopic pinnule much longer and 2 times width of acroscopic one; costae grooved above, continuous from rachis to costae; margins margins toothed, teeth spine-tipped.; veins free, forked.
Sori: round, in 1 row between midrib and margin, indusium: reniform, at a sinus, sporangia: brownish, maturity: midsummer.
Culture
Habitat: cool, moist woods at increasing elevation southward.
Distribution: eastern North America.
Hardy to -30°C, USDA Zone 4.
Synonyms
Aspidium campylopterum Kunze
Dryopteris austriaca (Jacquin) Schinz & Thellung
Dryopteris dilatata ssp. americana (Fisch.) Hulten
Dryopteris spinulosa (Swartz) Watt var. americana (Fischer ex Kunze) Fernald
Dryopteris spinulosa var. concordiana (Davenport) Eastman
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Dryopteris campyloptera.
Illustration from The Cultivated Species of the Fern Genus Dryopteris in the United States, Barbara Joe Hoshizaki and Kenneth A. Wilson, American Fern Journal, 89, 1, (1999), with permission.
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