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D. pacifica resources
Flora of JapanFlora of Japan
and frond photoand frond photo
Funabashi Herbarium, as D. varia var. hikonensisFunabashi Herbarium, as D. varia var. hikonensis
inoperativeMakino Herbarium, as Polystichum pacificum (link inoperative)
Hoshizaki and WilsonHoshizaki and Wilson
pair of frondspair of fronds
Photo index
sori, stipe and rachis scalessori, stipe and rachis scales

All Ferns
     aemula  affinis
     amurensis  arguta
     bissetiana  campyloptera
     carthusiana  caucasica
     celsa  championii
     clintoniana  crassirhizoma
     cristata  cycadina
     cystolepidota  decipiens
     dilatata  erythrosora
     expansa  filix-mas
     formosana  fragrans
     fuscipes  goldiana
     hondoensis  intermedia
     kuratae  lacera
     lepidopoda  ludoviciana
     marginalis  mindshelkensis
     oreades  pacifica
     polylepis  pseudo-filix-mas
     purpurella  remota
     sacrosancta  saxifraga
     sieboldii  stewartii
     sublacera  tokyoensis
     uniformis  varia

 Other Genera
    Adiantum  Arachniodes
    Aspidotis  Asplenium
    Astrolepis  Athyrium
    Blechnum  Cheilanthes
    Cryptogramma  Cyrtomium
    Cystopteris  Dennstaedtia
    Deparia  Diplazium
    Gymnocarpium  Lygodium
    Matteuccia  Onoclea
    Oreopteris  Osmunda
    Pellaea  Phegopteris
    Pleopeltis  Polypodium
    Polystichum  Pteridium
    Pteris  Pyrrosia
    Thelypteris  Woodsia
Dryopteris pacifica (Nakai) Tagawa
Etymology Pacifica is a reference to the area of the Pacific Ocean.
Description Rhizome: suberect to erect.
Frond: evergreen, monomorphic, blade/stipe ratio: 3:1 to 1:2.
Stipe: grooved, scales blackish-brown, vascular bundles: 3-7 in a c-shaped pattern.
Blade: 3-pinnate, triangular, abruptly narrowed tip, dark green, glossy, papery, narrow, black, somewhat bullate (inflated) scales.
Pinnae: 18 to 20 pair, alternate, strongly anadromic at the base, becoming catadromic above; pinnules downward-pointing, nearest the rachis, larger than upward ones; costae grooved above, continuous from rachis to costae; segments dentate; margins margins entire; veins free, forked.
Sori: round, small, in 1 row between midrib and margin, indusium: reniform, at a sinus, sporangia: brownish.
Culture Habitat: lowland forest edge. Distribution: Japan, Korea, China. Hardy to -20°C, USDA Zone 6.
Distinctive Characteristics The abruptly narrowed blade above the 10th or 11th set of pinnae is rare, but not unique; D. cystolepidota (nipponensis) has this, too, but here the lower pinnules closest to the rachis are longer.
Polystichum pacificum Nakai
Polystichum hoolepis var. hikonensis H. Itô
Dryopteris varia (L.) Kuntze var. hikonensis (H. Itô) Sa. Kurata
Dryopteris hikonensis (H. Ito) Nakaike
Compare to This shares several characteristics with D. cystolepidota, most notably the contracted blade tip, the 3-pinnate division at the base, toothed segment margins, and dark, blackish scales on stipe and rachis. D. cystolepidota, however, differs in having the second downward-pointing pinnules of the lower pinnae longest, whereas in D. pacifica has the ones closest to the rachis longest.
Compare to Similar to D. varia, differing from it in: scales at lower portion of stipes blackish brown to black; upper pinnae rather suddenly shortened to form acuminate apex; stiff-papery, shining on the upper surface; lobes minutely toothed at margin; indusia sometimes hairy at margin. Also similar to D. bissetiana, differing from it in the margins are not reflexed.
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