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D. pacifica resources
Flora
Flora of JapanFlora of Japan
Herbarium
and frond photoand frond photo
Funabashi Herbarium, as D. varia var. hikonensisFunabashi Herbarium, as D. varia var. hikonensis
inoperativeMakino Herbarium, as Polystichum pacificum (link inoperative)
Monograph
Hoshizaki and WilsonHoshizaki and Wilson
Photo
frondfrond
pair of frondspair of fronds
Photo index
sori, stipe and rachis scalessori, stipe and rachis scales

All Ferns
Dryopteridaceae
�� Dryopteris
���� aemula affinis
���� amurensis arguta
���� bissetiana campyloptera
���� carthusiana caucasica
���� celsa championii
���� clintoniana crassirhizoma
���� cristata cycadina
���� cystolepidota decipiens
���� dilatata erythrosora
���� expansa filix-mas
���� formosana fragrans
���� fuscipes goldiana
���� hondoensis intermedia
���� kuratae lacera
���� lepidopoda ludoviciana
���� marginalis mindshelkensis
���� oreades pacifica
���� polylepis pseudo-filix-mas
���� purpurella remota
���� sacrosancta saxifraga
���� sieboldii stewartii
���� sublacera tokyoensis
���� uniformis varia
���� wallichiana

�Other Genera
��� Adiantum Arachniodes
��� Aspidotis Asplenium
��� Astrolepis Athyrium
��� Blechnum Cheilanthes
��� Cryptogramma Cyrtomium
��� Cystopteris Dennstaedtia
��� Deparia Diplazium
��� Gymnocarpium Lygodium
��� Matteuccia Onoclea
��� Oreopteris Osmunda
��� Pellaea Phegopteris
��� Pleopeltis Polypodium
��� Polystichum Pteridium
��� Pteris Pyrrosia
��� Thelypteris Woodsia
��� Woodwardia
Dryopteris pacifica (Nakai) Tagawa
Etymology Pacifica is a reference to the area of the Pacific Ocean.
Description Rhizome: suberect to erect.
Frond: evergreen, monomorphic, blade/stipe ratio: 3:1 to 1:2.
Stipe: grooved, scales blackish-brown, vascular bundles: 3-7 in a c-shaped pattern.
Blade: 3-pinnate, triangular, abruptly narrowed tip, dark green, glossy, papery, narrow, black, somewhat bullate (inflated) scales.
Pinnae: 18 to 20 pair, alternate, strongly anadromic at the base, becoming catadromic above; pinnules downward-pointing, nearest the rachis, larger than upward ones; costae grooved above, continuous from rachis to costae; segments dentate; margins margins entire; veins free, forked.
Sori: round, small, in 1 row between midrib and margin, indusium: reniform, at a sinus, sporangia: brownish.
Culture Habitat: lowland forest edge. Distribution: Japan, Korea, China. Hardy to -20�C, USDA Zone 6.
Distinctive Characteristics The abruptly narrowed blade above the 10th or 11th set of pinnae is rare, but not unique; D. cystolepidota (nipponensis) has this, too, but here the lower pinnules closest to the rachis are longer.
Synonyms
Polystichum pacificum Nakai
Polystichum hoolepis var. hikonensis H. It�
Dryopteris varia (L.) Kuntze var. hikonensis (H. It�) Sa. Kurata
Dryopteris hikonensis (H. Ito) Nakaike
Notes
Compare to This shares several characteristics with D. cystolepidota, most notably the contracted�blade tip, the 3-pinnate division at the base, toothed segment margins, and dark, blackish scales on stipe and rachis. D. cystolepidota, however, differs in having the second downward-pointing pinnules of the lower pinnae longest, whereas in D.�pacifica hasthe ones closest to the rachis longest.
Compare to Similar to D. varia, differing from it in: scales at lower portion of stipes blackish brown to black; upper pinnae rather suddenly shortened to form acuminate apex; stiff-papery, shining on�the upper�surface; lobes minutely toothed at margin; indusia sometimes hairy at margin. Also similar to D. bissetiana, differing from it in the margins are not reflexed.
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