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D. bissetiana resources
Flora
Flora of JapanFlora of Japan
Monograph
Hoshizaki and WilsonHoshizaki and Wilson
Photo
basal pinnae, close-up of incised veins, click link for the reversebasal pinnae, close-up of incised veins, click link for the reverse
habithabit
pair of frondspair of fronds

All Ferns
Dryopteridaceae
�� Dryopteris
���� aemula affinis
���� amurensis arguta
���� bissetiana campyloptera
���� carthusiana caucasica
���� celsa championii
���� clintoniana crassirhizoma
���� cristata cycadina
���� cystolepidota decipiens
���� dilatata erythrosora
���� expansa filix-mas
���� formosana fragrans
���� fuscipes goldiana
���� hondoensis intermedia
���� kuratae lacera
���� lepidopoda ludoviciana
���� marginalis mindshelkensis
���� oreades pacifica
���� polylepis pseudo-filix-mas
���� purpurella remota
���� sacrosancta saxifraga
���� sieboldii stewartii
���� sublacera tokyoensis
���� uniformis varia
���� wallichiana

�Other Genera
��� Adiantum Arachniodes
��� Aspidotis Asplenium
��� Astrolepis Athyrium
��� Blechnum Cheilanthes
��� Cryptogramma Cyrtomium
��� Cystopteris Dennstaedtia
��� Deparia Diplazium
��� Gymnocarpium Lygodium
��� Matteuccia Onoclea
��� Oreopteris Osmunda
��� Pellaea Phegopteris
��� Pleopeltis Polypodium
��� Polystichum Pteridium
��� Pteris Pyrrosia
��� Thelypteris Woodsia
��� Woodwardia
Dryopteris bissetiana (Baker) C. Chr.

Beaded wood fern

Etymology For a collector in Miyanosh'ta, Japan, Mr. Bissett, otherwise unidentified by J.G. Baker.
Description Rhizome: short-creeping, clumping, scaly.
Frond: 45 cm high by 30 cm wide, evergreen, monomorphic, blade/stipe ratio: 3:1 to 1:2.
Stipe: grooved, scales black, narrow triangular, vascular bundles: 3-7 in a c-shaped pattern.
Blade: 3-pinnate only at the base, deltate-ovate to lanceolate, leathery, embossed upper surface, costa with small bullate scales below.
Pinnae: 15 to 20 pair, lowest pinnae anadromous; pinnules triangular, curved outwards, almost entire above to pinnate below, the first lower pinnule of the lowest pinnae twice the lenth of the first upper pinnule; costae grooved above, continuous from rachis to costae; margins serrate, slightly reflexed; veins free, forked, visible on both surfaces.
Sori: round, submarginal to medial, ultimate segments each bearing one sorus, indusium: reniform, greenish at maturity, attached at a sinus, sporangia: brownish, maturity: fall.
Culture Habitat: forest floor, edge of mountain forests. Distribution: Japan, Korea, China. Hardy to -25�C, USDA Zone 5.
Synonyms
Nephrodium bissetianum Baker
Polypodium setosum Thunb
Dryopteris varia (L.) Kuntze var. setosa (Thunb.) Ohwi
Polystichum bissetianum Nakai
Polystichum sacrosanctum Koidzumi, misapplied
Dryopteris bissetiana
Dryopteris bissetiana. Frond; the embossed appearance (seen more clearly in the click-through enlargement) of the upper surface is distinctive. �Photo: Tom Stuart
Dryopteris bissetiana
Dryopteris bissetiana. Pinna; click through for a blowup of the center. �Photo: Tom Stuart
Dryopteris bissetiana
Dryopteris bissetiana. �Illustration from The Cultivated Species of the Fern Genus Dryopteris in the United States, Barbara Joe Hoshizaki and Kenneth A. Wilson, American Fern Journal, 89, 1, (1999), with permission.
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