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D. mindshelkensis resources
Ferns of the British Isles, as D. submontanaFerns of the British Isles, as D. submontana
frond (as D. villarii ssp pallida)frond (as D. villarii ssp pallida)
Hoshizaki and WilsonHoshizaki and Wilson
inoperativehabit (as D. villarii) (link inoperative)

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Dryopteris mindshelkensis Pavlov

Rigid buckler fern, limestone wood fern

Etymology For Mindshelke, a district in the Karatau Mountains of southern Kazakhstan and the type location.
Description Rhizome: short-creeping, forming clumps of shuttlecocks.
Frond: 60 cm high by 15 cm wide, deciduous, monomorphic, blade/stipe ratio: 3:2.
Stipe: grooved, pale brown, vascular bundles: 3-7 in a c-shaped pattern.
Blade: 2-pinnate, deltate-ovate to lanceolate, stiff, somewhat leathery, distinctly mealy, grayish-green, covered with glands.
Pinnae: 12 to 20 pair, lanceolate, often slightly ascending; costae grooved above, continuous from rachis to costae; margins serrate, spiny; veins free, forked.
Sori: round, in 1 row between midrib and margin, indusium: reniform, pale green when young, at a sinus, sporangia: brownish, maturity: summer.
Culture Habitat: among limestone rocks. Distribution: Spain, Algeria, east to the Caucasus and northern Iran and Kazakhstan. Hardy to -20C, USDA Zone 6.
Dryopteris submontana (Fraser-Jenk. & Jermy) Fraser-Jenk.
Dryopteris villarii (Bell.) Woyn. ex Shinz. & Thell. ssp. submontana Fraser-Jenk. & Jermy
Lastrea rigida (Sw.) C. Presl., misapplied
Dryopteris villarii ssp. mindshelkensis (Pavlov) Fraser-Jenk.
Dryopteris mindshelkensis
Dryopteris mindshelkensis.  Illustration from The Cultivated Species of the Fern Genus Dryopteris in the United States, Barbara Joe Hoshizaki and Kenneth A. Wilson, American Fern Journal, 89, 1, (1999), with permission.
Dryopteris mindshelkensis
Dryopteris mindshelkensis. Frond, stipe indusium, pinnule. As Lastrea rigida, one of many names for this species.  Drawing by J.E.S. Fecit
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