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P. kruckebergii resources
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All Ferns
     acrostichoides  aculeatum
     andersonii  braunii
     craspedosorum  imbricans
     kruckebergii  lemmonii
     lepidocaulon  lonchitis
     makinoi  munitum
     neolobatum  polyblepharum
     proliferum  retroso-paleaceum
     rigens  scopulinum
     setiferum  setigerum
     tagawanum  tripteron
     tsus-simense  xiphophyllum

 Other Genera
    Adiantum  Arachniodes
    Aspidotis  Asplenium
    Astrolepis  Athyrium
    Blechnum  Cheilanthes
    Cryptogramma  Cyrtomium
    Cystopteris  Dennstaedtia
    Deparia  Diplazium
    Dryopteris  Gymnocarpium
    Lygodium  Matteuccia
    Onoclea  Oreopteris
    Osmunda  Pellaea
    Phegopteris  Pleopeltis
    Polypodium  Pteridium
    Pteris  Pyrrosia
    Thelypteris  Woodsia
Polystichum kruckebergii W. H. Wagner

Kruckeberg's holly fern

Etymology Named after botanist and plant ecologist Dr. Arthur Kruckeberg (1920- )
Description Rhizome: erect, short, sparsely scaly; scales light brown, gradually diminishing in size upwards.
Frond: 35 cm high by 3 cm wide, evergreen, monomorphic or weakly dimorphic, blade/stipe ratio: 4:1 to more usually 10:1.
Stipe: grooved, sparesly scaly, vascular bundles: 4 or more, in an arc.
Blade: 1-pinnate, lobed pinnae, sometimes pinnatifid, lanceolate, base narrowed, glossy, microscales lanceolate with few projections, confined to costa below.
Pinnae: 20 to 40 pair, rhombic-ovate, 0.5--1.5 cm, lower ones lobed at the base, the upper ear well-developed; margins toothed, spiny; veins free, forked.
Sori: round, undersides of the upper pinnae, indusium: peltate, central, sporangia: dark brown, maturity: midsummer.
Dimensionality: pinnae twisted somewhat out of plane of blade.
Culture Habitat: rocks and cliffs in subalpine to alpine habitats, 21003200 m. in California, lower elevations north. Distribution: western North America. Hardy to -20C, USDA Zone 6.
Polystichum kruckebergii
Polystichum kruckebergii. a) fronds, only upper half fertile, confluent at maturity; b) part of fertile pinna.  Illustration by V. Fulford from Ferns and Fern Allies of Canada, William J. Cody and Donald M. Britton, 1989, Agriculture Canada, used with permission.
Compare to distinguished from the common P. lonchitis by the lobed basal pinnae, and from P. scopulinum by the spiny pinnae margins.
Parents probably P. lonchitis and P. lemmonii. The latter is clearly 2-pinnate.
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