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D. cystolepidota resources
Flora of Japan, as <i>D. nipponensis</i>Flora of Japan, as D. nipponensis
with live frondwith live frond
Hoshizaki and WilsonHoshizaki and Wilson
fronds, as <i>D. erythrosora v. dilatata</i>fronds, as D. erythrosora v. dilatata
inoperativesolitary frond (link inoperative)
sori, stipe scalessori, stipe scales

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     aemula  affinis
     amurensis  arguta
     bissetiana  campyloptera
     carthusiana  caucasica
     celsa  championii
     clintoniana  crassirhizoma
     cristata  cycadina
     cystolepidota  decipiens
     dilatata  erythrosora
     expansa  filix-mas
     formosana  fragrans
     fuscipes  goldiana
     hondoensis  intermedia
     kuratae  lacera
     lepidopoda  ludoviciana
     marginalis  mindshelkensis
     oreades  pacifica
     polylepis  pseudo-filix-mas
     purpurella  remota
     sacrosancta  saxifraga
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     sublacera  tokyoensis
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    Adiantum  Arachniodes
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    Blechnum  Cheilanthes
    Cryptogramma  Cyrtomium
    Cystopteris  Dennstaedtia
    Deparia  Diplazium
    Gymnocarpium  Lygodium
    Matteuccia  Onoclea
    Oreopteris  Osmunda
    Pellaea  Phegopteris
    Pleopeltis  Polypodium
    Polystichum  Pteridium
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    Thelypteris  Woodsia
Dryopteris cystolepidota (Miq.) C. Chr.
Etymology Greek cysto bladder + lepis scale, referring to the bullate (puffed base) scales.
Description Rhizome: short-creeping.
Frond: 60 cm high by 25 cm wide, evergreen, monomorphic, blade/stipe ratio: 2:1.
Stipe: grooved, scales dull brown to black, to 10mm, vascular bundles: 3-7 in a c-shaped pattern.
Blade: 3-pinnate at base, triangular, abruptly narrowed tip, thin leathery, slightly glossy, new growth red, costae with bullate scales below.
Pinnae: 15 to 20 pair, lowest pair longest; pinnules larger ones eared on both sides, generally quite long, the second downward-pointing ones on the lower pinnae the longest; costae grooved above, continuous from rachis to costae; veins free, forked.
Sori: round, medial, indusium: reniform, pale green early (reports of reddish tints are probably erroneous), at a sinus, sporangia: brownish.
Culture Habitat: on forest floor in lowlands . Distribution: Japan and Korea. Hardy to -20C, USDA Zone 6.
Distinctive Characteristics Flora of Japan: "similar to D. erythrosora but different in: scales on stipes blackish brown, those on rachis and pinna rachis also blackish brown, rather dense, caducous; laminae broadly ovate, abruptly acuminate at apex, basal basiscopic pinnules of lowest pinnae larger, other pinnules oblong-ovate to ovate, shallowly crenate to pinnatifid, lower ones stalked; tenderly papyraceous, not shining; sori smaller; indusia not tinted reddish."
Dryopteris nipponensis Koidz.
Dryopteris erythrosora (D. C. Eaton) Kuntze var. dilatata (Koidz.) Sugim.
Dryopteris nipponensis Koidz. var. dilatatum Koidz.
Aspidium cystolepidotum Miq.
Dryopteris cystolepidota
Dryopteris cystolepidota.  Illustration from The Cultivated Species of the Fern Genus Dryopteris in the United States, Barbara Joe Hoshizaki and Kenneth A. Wilson, American Fern Journal, 89, 1, (1999), with permission.
Compare to This shares several characteristics with D. pacifica, most notably the contracted blade tip, the 3-pinnate division at the base, toothed segment margins, and dark, blackish scales on stipe and rachis. This differs, however, in having the second downward-pointing pinnules of the lower pinnae longest, whereas in D. pacifica the ones closest to the rachis are longest.
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