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C. montana resources
Distribution
North AmericaNorth America
WorldWorld
Drawing
Britton & Brown, as Filix montanaBritton & Brown, as Filix montana
Illustrations of the British Flora, 1924, Walter Hood FitchIllustrations of the British Flora, 1924, Walter Hood Fitch
Flora
Flora of North AmericaFlora of North America
Photo
fertile frondfertile frond
frondfrond
frond, habitfrond, habit
Photo index
CalPhotosCalPhotos
habitat, fronds, sorihabitat, fronds, sori

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Cystopteris montana (Lamarck) Bernhardi ex Desvaux

Mountain bladder fern

Etymology Montana is the word for the mountains.
Description Rhizome: long-creeping, cordlike, internodes 1--2 cm, whitish at tip, scales usually tan to light brown, ovate-lanceolate.
Frond: 40 cm high by 15 cm wide, deciduous, monomorphic, blade/stipe ratio: 1:3 to 1:1.
Stipe: dark brown to black at base, gradually becoming green or straw-colored above, sparsely scaly throughout, vascular bundles: 2, round or oblong.
Blade: 3-pinnate-pinnatifid or more, equilaterally triangular, similar to a Gymnocarpium in form, firm, translucent, pale green, rachis and costae with tan, multicellular gland-tipped hairs.
Pinnae: 7 to 8 pair, anadromous, pinnae often bending and/or curving towards the tip of the frond; pinnules lower innermost pinnule of the lowest pinnae large, resembling perhaps the third pinna above; costae grooves above continuous from rachis to costae; margins serrate; veins free, simple or forked, directed to notches.
Sori: round, in 1 row between midrib and margin, indusium: ovate, transparent, beneath sorus on midrib side, sporangia: black, maturity: midsummer to late summer.
Dimensionality: just above the basal pinna pair, the rachis curves over to hold the blade horizontally.
Culture Habitat: calcareous wet woods or along water courses, or in subalpine-arctic Salix communities. Distribution: boreal or subalpine Northern Hemisphere, never abundant. Hardy to -40�C, USDA Zone 2, requires a cool summer.
Distinctive Characteristics Elegant is a term that pops up in many descriptions. Certainly very lacy, evocative of Gymnocarpium in blade form and horizontal position. The only not-very-nice thing about it is the poisonous hydrocyannic acid emitted by bruised fronds. Perhaps not in cultivation, but impossible to omit.
Synonyms
Polypodium montanum Lamarck
Cystopteris montana
Cystopteris montana. top:segment with sori and glandular hairs on costa, left: stips scales, right: two pinnules�closest to rachis. �Illustration from Scandinavian Ferns by Benjamin �llgaard and Kirsten Tind, Rhodos, 1993.
Notes
Compare to There are a number of thrice+ divided, small ferns, but all are of a clumping habit. Gymnocarpium species are not quite as divided, but do possess the running habit and dimensionality of this plant; Gymnocarpium, however, has black stipes, lacks indusia.
Cystopteris montana
Cystopteris montana. a) frond, highly dissected; b) fertile pinnule or segment with veins ending in the notches. �Illustration by V. Fulford from Ferns and Fern Allies of Canada, William J. Cody and Donald M. Britton, 1989, � Agriculture Canada, used with permission.
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